Using gland regions on their heads, warthogs mark poles and other objects for self-orientation. Males often mark during the mating season. The male also sprays urine when searching for and inspecting sleeping cavities.
1965
1933
2023
2005
2018
2014
2015
2012
2013
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1997
1996
2019
2017
1937
1990
2004
2006